This is a brief overview of some of the literature dealing with flying snakes. I've tried to be as comprehensive as possible. If you come across other articles not listed, please let me know.

      --Popular Articles--

A more updated list can be found on the Press page of the Socha Lab web site.

Snakes Take Flight. By Cynthia Wei. Journal of Experimental Biology Inside JEB piece, May 15 2005. About the two flying snake research papers published in issue 208(10) (listed below). |pdf|

Serpents in the Air. By Adam Summers. Natural History magazine, May 2003. About flying snake research; includes a nice illustration.

Wild Gliders. By Tim Laman. National Geographic magazine, October 2000, pages 68-85. Amazing and unique photographs of gliders in Borneo.

Flying Snakes. By Sharon Parmet. MUSE magazine Volume 3, Number 7, September 1999, pages 35-38. An article for kids.

Flyings Snakes. By Christopher Miller. Popular Science magazine, February 1999, page 26. A short blurb.

Gliders and Parachuters. By John C. Murphy. Reptiles magazine, March 1996, pages 12-26. An interesting account of John's travels in Borneo discussing flying frogs, snakes, and lizards.

Notes on the Biology and Captive Care of the Flying Snakes (Chrysopelea). By Lenny Flank, Jr. Reptile and Amphibians Hobbyist magazine, July 1999, pages 28-32. Captive care, focusing on C. ornata.

      --Scientific Journal and Book Articles--

Yeaton, I.J., G. Baumgardner, S.D. Ross, and J.J. Socha. 2020. Undulation enables gliding in flying snakes. Nature Physics 51(6): 969-982. |link|

S Yeaton, I.J., J.J. Socha, and S.D. Ross. 2017. Global dynamics of non-equilibrium gliding in animals. Bioinspiration and Biomimetics 12(2): 026013. |pdf|

Jafari, F., S. Tahmasian, S.D. Ross, and J.J. Socha. 2017. Control of gliding in a flying snake-inspired n-chain model. Bioinspiration and Biomimetics 12(6) 066002. |pdf|

Socha, J.J., F. Jafari, Y. Munk, and G. Byrnes. 2015. How animals glide: from trajectory to morphology. Canadian Journal of Zoology 93: 901-924. |pdf|

Jafari, F., S.D. Ross, P.P. Vlachos, and J.J. Socha. 2014. A theoretical analysis of gliding in flying snakes. Bioinspiration and Biomimetics 9(2): 025014 |pdf|

Krishnan, A., J.J. Socha, P.P. Vlachos, and L.A. Barba. 2014. Lift and wakes of flying snakes. Physics of Fluids. 26: 031901. |pdf|

Holden, D., J.J. Socha, N.D. Cardwell, and P.P. Vlachos. 2014. Aerodynamics of the flying snake, Chrysopelea paradisi: how a bluff-body cross-sectional shape contributes to gliding performance. Journal of Experimental Biology 217 (3): 382-394. |pdf|

Socha, J.J.. 2011. Gliding flight in Chrysopelea: Turning a snake into a wing. Integrative and Comparative Biology 51(6): 969-982. |pdf|

Socha, J.J., K. Miklasz, F. Jafari, P.P. Vlachos. 2010. Non-equilibrium trajectory dynamics and the kinematics of gliding in a flying snake. Bioinspiration and Biomimetics 5 (2010) 045002. |pdf|

Miklasz, K., M. LaBarbera, X. Chen, J.J. Socha. 2010. Effects of body cross-sectional shape on flying snake aerodynamics. Experimental Mechanics 50(9): 1335-1348. |pdf|

Socha, J.J., T. O'Dempsey, M. LaBarbera. 2005. A three-dimensional kinematic analysis of gliding in a flying snake, Chrysopelea paradisi. Journal of Experimental Biology 208(10): 1817-1833. |pdf|

Socha, J.J. and M. LaBarbera. 2005. Effects of size and behavior on aerial performance of two species of flying snakes (Chrysopelea). Journal of Experimental Biology 208(10): 1835-1847. |pdf|

Socha, J.J. and C.A. Sidor. 2005. Chrysopelea ornata, C. paradisi (Flying Snakes). Behavior. Herpetological Review 36(2): 190-191. |pdf|

Socha, J.J. 2002a. Gliding flight in the paradise tree snake. Nature 418, 603-604. |pdf|

Socha, J.J. 2002b. The Biomechanics of Flight in Snakes. Ph.D. Thesis, University of Chicago. 262 pp. |pdf| **note--16.1 mb file

Socha, J.J., and T. O'Dempsey. (2001). A three-dimensional analysis of flying snake flight. American Zoologist (5):390A. (abstract) |pdf|

Lim, K.P. and L.J. Lim (1999). The terrestrial herpetofauna of Pulau Tioman, peninsular Malaysia. The Raffles Bulletin of Zoology Supplement 6, 131-155.

Steubing, R.B. and R.F. Inger (1999). A Field Guide to the Snakes of Borneo. Borneo: Natural History Publications. 254 pp. Amazon link

Socha, J.J. (1999). The take-off and landing kinematics of a flying snake, Chrysopelea ornata. American Zoologist 39(5):104A-105A. (abstract) |pdf|

Socha, J.J. (1998). A description of 'flight' kinematics in the flying snake, Chrysopelea paradisi. American Zoologist 38(5):150A. (abstract) |pdf|

Manthey, U. and W. Grossman (1997). Amphibien and Reptilien Sudostasiens. Berlin, Verlag.

Dowling, H. G., C. A. Hass, S. B. Hedges and R. Highton (1996). Snake relationships revealed by slow-evolving proteins: a preliminary survey. Journal of Zoology, London 240, 1-28.

Murthy, T. S. N. (1990). Illustrated guide to the snakes of the Western Ghats, India. Records of the Zoological Survey of India 114, 1-76.

Rawlinson, P. A. (1990). The terrestrial vertebrate fauna of the Krakatau Islands Sunda Strait Indonesia 1883-1986. Philosophical Transactions of the Royal Society of London Series B 328(1245), 3-28.

Basu, D. (1989). Range extension of Chrysopelea ornata Shaw (Reptilia: Colubridae) with comments on the distribution of some snakes in north India. Journal of Bombay Natural History Society 86(1), 110-111.

Malkmus, R. (1987). Herpetological observations on Mount Kinabalu Borneo. Mitteilungen aus dem Zoologischen Museum in Berlin 63(2), 269-262.

Gaulke, M. (1986). Zum beutefangverhalten der schmuckbaumschlange Chrysopelea paradisi Boie, 1827. Salamandra 22(2/3), 211-213.

Tweedie, M. W. F. (1983). The Snakes of Malaya. Singapore, Singapore National Printers. 65 pp.

de Silva, P. (1980). Snake Fauna of Sri Lanka. Sri Lanka, National Museums of Sri Lanka. 472 pp.

Mundkur, T. (1978). A note on reproduction in the flying snake Chrysopelea ornata (Shaw). J. Bombay Nat. Hist. Soc. 75(3), 854-859.

Heyer, W. R. and S. Pongsapipatana (1970). Gliding speeds of Ptychozoon lionatum (Reptilia: Gekkonidae) and Chrysopelea ornata (Reptilia: Colubridae). Herpetologica 26, 317-319.

Mertens (1970). Zur Frage der "Fluganpassungen" von Chrysopelea (Serpentes, Colubridae). Salamandra 6, 11-14. [English translation: On the question of "flight adaptation" in Chrysopelea (Serpentes, Colubridae). Bulletin of the Chicago Herpetological Society 21(1-2), 39-41.]

Mertens, R. (1968). Die arten und unterarten der schmuckbaumschlangen (Chrysopelea). Senckenbergiana biologica 49(3/4), 191-217.

Leviton, A. E. Contributions to a review of Philippine snakes, IV: The genera Chrysopelea and Dryophiops. Philippine Journal of Science 93, 131-145.

Gharpurey, K. (1962). Snakes of India and Pakistan. Bombay, Popular Prakashan. 89 pp.

Mertens, R. (1960). Fallschirmspringer und Gleitflieger unter den Amphibien und Reptilien. Der Flug der Tiere. Frankfurt, Waldemar Kramer Verlag: 135-144. [English translation: Gliding and parachuting flight among the amphibians and reptiles. Bulletin of the Chicago Herpetological Society 21(1-2), 42-46.]

Tweedie, M. W. F. (1960). The Malayan gliding reptiles. Proc. South London Entomol. and Nat. Hist. Soc. 1959, 97-103.

Vaughn-Arbuckle, K. H. (1959). Flying snakes. Journal of the Bombay Natural History Society 56(3), 640-642.

Tweedie, M. W. F. (1954). Notes on Malayan Reptiles, No. 3. Bulletin of the Raffles Museum 25, 107-117.

Oliver, J. A. (1951). "Gliding" in amphibians and reptiles, with a remark on an arboreal adaptation in the lizard, Anolis carolinensis carolinensis Voigt. The American Naturalist 85, 171-176.

Constable, J. D. (1949). Reptiles from the Indian peninsula in the museum of comparative zoology. Bulletin of the Museum of Comparative Zoology 103(2).

Shebbeare, E. O. (1941). "Flying" snakes, tree shrews, and mimicry. Malayan Nat. Journal 1, 157-162.

Hediger, H. (1932). Zum problem der "fliegenden" schlangen. Revue Suisse de zoologie 39(5), 239-246.

Pendlebury, H. M. (1931). A "flying" snake. Bulletin of the Raffles Museum 5, 75. Wall, F. (1908). A popular treatise on the common Indian snakes, Part 6. Journal of the Bombay Natural History Society 18(2), 227-243.

Shelford, M. A. (1906). A note on "flying" snakes. Proc. Zool. Soc. Lond. 1(16), 227-230.

Daly, M. (1899). A flying snake. Journal of the Bombay Natural History Society XII, 589.

Flower, S. S. (1899). Notes on a second collection of reptiles made in the Malay Peninsula and Siam, from November 1896 to September 1898, with a list of the species recorded from those countries. Proceedings of the Zoological Society of London May 16, 682-685.

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